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It is intriguing that 4-HNE, as a signaling molecule, can regulate the expression of many transcription factors while enhancing the antioxidant mechanism of cells

170 In periosteum-derived skeletal progenitors, pharmacological inhibition of Complex III of the electron transport chain increased succinate and 2-hydroxyglutarate levels that inhibited ten-eleven translocation (TET) DNA demethylase activity, causing wide-spread DNA hypermethylation in the genome

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doi: 10.1371/journal.pone.0014356